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Research

Since a complete list of my publications can be found on the NASA/SAO ADS as well as my CV, I’ll organize this page by research topic rather than date or number of citations. Papers on which I am an author are bolded, and the bibliography at the bottom of this page links to the papers themselves. This is not a full review, so not all relevant papers are cited; for that I’d recommend traversing the citation graph on the ADS.

Giant Planet Compositions

Mass-metallicity relation

Giant exoplanets with equilibrium temperatures below 1000 K have radii that are well explained by the same interior structure models that were developed for Jupiter and Saturn. As such, we can infer their bulk compositions by adjusting the parameter in structure models such that radius matches its observed value. I did this for 47 giant planets, finding that the bulk metal mass was related to the mass as approximately MzM0.61M_z \propto M^{0.61} (Thorngren et al. 2016). Since the bulk metallicity is the mass fraction Z=Mz/MZ=M_z/M, this means metallicity is negatively correlated by mass.

This scaling resembles that of mass of solids in the protoplanetary disk within the gravitational influence of the planet during formation, suggesting that this late accretion of solids plays an important role in determining final compositions. It also implies that these planets will typically have low C/O ratios in their atmospheres (Espinoza et al. 2017). While core-accretion models might suggest linear relationship between stellar and planetary metallicities, comparisons with stellar metallicities indicate the relationship is much weaker than that (Teske et al. 2019; Thorngren et al. 2021), perhaps as a result of the chaotic nature of planet formation.

Bulk compositions also have value in support of atmospheric observations, especially transmission and emission spectroscopy. The atmospheric metallicity cannot exceed the bulk metallicity stably (Thorngren and Fortney 2019), allowing the latter to serve as an easily-obtained upper-limit on the plausible atmospheric metallicity (Kreidberg et al. 2018; Mikal-Evans et al. 2021; Baxter et al. 2021; Bean et al. 2023).

With the successful launch and commissioning of the James Webb Space Telescope (JWST), we have been able to go further. If an upper-limit on the atmospheric metallicity can be established, then knowing the bulk metallicity sets a lower limit on the core mass of the planet (Sing et al. 2024), including any “fuzzy” core layers (Bloot et al. 2023). This is an exciting new frontier in exoplanet science, as the core mass corresponds to the mass during formation at which the planet was able to retain large quantities of H/He, beginning runaway accretion (Helled and Stevenson 2017).

Hot Jupiter Heating

Flux-radius diagram.

Hot Jupiters, defined for this purpose as those with equilibrium temperatures above 1000 K, are unexpectedly large compared to our models. My review chapter, Thorngren (2024) (figure right), covers some of the history and current state of this problem. While we have made a great deal of progress in understanding the effect, the problem has not yet been resolved.

A key part of the challenge is that for these planets, both the internal temperatures and the composition are not known, preventing us from inferring either directly from the radius. My contribution to this area was Thorngren and Fortney (2018), in which I used a hierarchical Bayesian model that identified the internal temperatures that would correspond to these planets having the same distribution of compositions seen for the warm giants (Thorngren et al. 2016). Modeling this as a single source of additional heat deep within the planet, we found that it peaks as a fraction if the incoming light energy at Teq1600T_\mathrm{eq} \approx 1600 K. This matches predictions for the Ohmic dissipation explanation (Menou 2012), as a result of a strong dependence on the atmospheric wind speeds, which feature that same peak. However, it could also support other processes that depend on the wind speed, like temperature advection (Tremblin et al. 2017).

This heating has important implications for other aspects of hot Jupiter physics. First, it suggests that their intrinsic temperatures (a measure of heat rising from the interior) is much higher (Thorngren et al. 2019) than the 100 K of Jupiter, which in turn pushes their radiative convective boundaries to higher altitudes / lower pressures. This is an important input for global circulation models (Komacek et al. 2022), and affects the pressure-temperature profile, cloud structure (Gao et al. 2020), and disequilibrium chemistry (Fortney et al. 2020) as well. Finally, a larger intrinsic flux could allow for a very strong magnetic field (Yadav and Thorngren 2017), though this would depend on the location and nature of the magnetic dynamo.

The age-dependence of hot Jupiter radii, can tell us about the process that heats them as well: (Lopez and Fortney 2016) observe that when TeqT_\mathrm{eq} increases, whether the planet grows larger (reinflates) depends on whether the effect is heat trapping or active heating of the interior. A number of papers have followed up on this; for my part, I observed in (Thorngren et al. 2021) that hot Jupiters orbiting main-sequence stars show evidence of both heat trapping and active heating.

Mass Loss and Tidal Evolution

Tides and XUV-driven mass loss were a natural area for my research to branch out in, as they are both driven by the radius and affect the interior evolution of the planet. For example, XUV-driven mass loss rates scale as the radius cubed, but losing mass can change the radius of the planet. In Thorngren et al. (2023) I found that this results in a feedback loops for hot Saturns which, under the right conditions, can remove large portions of a planet’s gaseous envelope.

This experience, along with my more standard interior structure models, has allowed me to contribute to a number of projects on the fascinating case of super-puffs (Yoshida et al. 2023; Vissapragada et al. 2024; Thao et al. 2024; Karalis et al. 2025; Yee et al. 2025). These planets have both puzzlingly large radii for their low masses (hence the name) and appear to push the boundaries of gaseous planet stability. It remains unclear whether this is due to low metallicity or high temperatures, as either result is surprising.

Tidal evolution is related in a couple of ways. First, the mass loss process itself can push a planet into higher orbits (Valsecchi et al. 2014), which we considered in Thorngren et al. (2023). Second, eccentric planets can undergo tidal circularization, which reduces the semimajor axis and eccentricity while heating the planetary interior, enlarging it. This may help to explain some warm, low-density planets (Morley et al. 2017; Dang et al. 2022; Piaulet et al. 2023; Sing et al. 2024). Finally, in the most extreme cases (Wahl et al. 2021), the tides of a star on a planet can pull it into a prolate ellipsoid shape (somewhat like a football or egg).

Astrostatistics

While many of the aforementioned projects have had a strong statistical component (Thorngren and Fortney 2018; Thorngren et al. 2021), there are others for which the statistics were my primary contribution. Movshovitz et al. (2020) explored the space of Saturn interior structures consistent with the data, while not incorporating any equations of state a priori; my involvement was mainly helping to properly sampling the quite challenging posterior distribution. In Mayorga et al. (2020), I helped to design the statistical model and wrote the core code used to fit them to the data.

Sigma Comparison

More recently, in Thorngren et al. (2025) I observed that a statistical technique widely used in exoplanet atmospheres was invalid. The error was based on a misunderstanding of Sellke et al. (2001), which gave a way to interpret the p-values of simple hypotheses as an upper limit on the Bayes factor. Astronomers were using this to obtain an upper limit on the p-value from their Bayes factors, but this violated the requirements of the original formula and generally overestimates the confidence compared to proper significance tests. The figure on the right shows a comparison; the erroneous method is in orange and the standard Bayesian approach is in blue. I also discussed the prior sensitivity of Bayes factor analyses, and suggest the use of the simplified Bayesian predictive information criterion (BPICS, Ando (2011)) as a less prior-sensitive supporting test.


Ando, Tomohiro. 2011. “Predictive Bayesian Model Selection.” American Journal of Mathematical and Management Sciences 31 (1-2): 13–38. https://doi.org/10.1080/01966324.2011.10737798.
Baxter, Claire, Jean-Michel Désert, Shang-Min Tsai, et al. 2021. “Evidence for Disequilibrium Chemistry from Vertical Mixing in Hot Jupiter Atmospheres: A Comprehensive Survey of Transiting Close-in Gas Giant Exoplanets with Warm- Spitzer /IRAC.” Astronomy & Astrophysics 648 (April): A127. https://doi.org/10.1051/0004-6361/202039708.
Bean, Jacob L., Qiao Xue, Prune C. August, et al. 2023. “High Atmospheric Metal Enrichment for a Saturn-mass Planet.” Nature 618 (June): 43–46. https://doi.org/10.1038/s41586-023-05984-y.
Bloot, S., Y. Miguel, M. Bazot, and S. Howard. 2023. “Exoplanet Interior Retrievals: Core Masses and Metallicities from Atmospheric Abundances.” Monthly Notices of the Royal Astronomical Society 523 (August): 6282–92. https://doi.org/10.1093/mnras/stad1873.
Dang, Lisa, Taylor J. Bell, Nicolas B. Cowan, et al. 2022. “Thermal Phase Curves of XO-3b: An Eccentric Hot Jupiter at the Deuterium Burning Limit.” The Astronomical Journal 163 (January): 32. https://doi.org/10.3847/1538-3881/ac365f.
Espinoza, Néstor, Jonathan J. Fortney, Yamila Miguel, Daniel Thorngren, and Ruth Murray-Clay. 2017. “Metal Enrichment Leads to Low Atmospheric C/O Ratios in Transiting Giant Exoplanets.” The Astrophysical Journal Letters 838 (March): L9. https://doi.org/10.3847/2041-8213/aa65ca.
Fortney, Jonathan J., Channon Visscher, Mark S. Marley, et al. 2020. “Beyond Equilibrium Temperature: How the Atmosphere/Interior Connection Affects the Onset of Methane, Ammonia, and Clouds in Warm Transiting Giant Planets.” The Astronomical Journal 160 (December): 288. https://doi.org/10.3847/1538-3881/abc5bd.
Gao, Peter, Daniel P. Thorngren, Elspeth K. H. Lee, et al. 2020. “Aerosol Composition of Hot Giant Exoplanets Dominated by Silicates and Hydrocarbon Hazes.” Nature Astronomy 4 (May): 951–56. https://doi.org/10.1038/s41550-020-1114-3.
Helled, Ravit, and David Stevenson. 2017. “The Fuzziness of Giant PlanetsCores.” The Astrophysical Journal 840 (May): L4. https://doi.org/10.3847/2041-8213/aa6d08.
Karalis, Amalia, Eve J. Lee, and Daniel P. Thorngren. 2025. “Separating Super-puffs Versus Hot Jupiters Among Young Puffy Planets.” The Astrophysical Journal 978 (January): 46. https://doi.org/10.3847/1538-4357/ad946c.
Komacek, Thaddeus D., Peter Gao, Daniel P. Thorngren, Erin M. May, and Xianyu Tan. 2022. “The Effect of Interior Heat Flux on the Atmospheric Circulation of Hot and Ultra-hot Jupiters.” The Astrophysical Journal Letters 941 (2): L40. https://doi.org/10.3847/2041-8213/aca975.
Kreidberg, Laura, Michael R. Line, Daniel Thorngren, Caroline V. Morley, and Kevin B. Stevenson. 2018. “Water, High-altitude Condensates, and Possible Methane Depletion in the Atmosphere of the Warm Super-Neptune WASP-107b.” The Astrophysical Journal Letters 858 (May): L6. https://doi.org/10.3847/2041-8213/aabfce.
Lopez, Eric D., and Jonathan J. Fortney. 2016. “Re-Inflated Warm Jupiters Around Red Giants.” The Astrophysical Journal 818 (February): 4. https://doi.org/10.3847/0004-637X/818/1/4.
Mayorga, L. C., David Charbonneau, and D. P. Thorngren. 2020. “Reflected Light Observations of the Galilean Satellites from Cassini: A Test Bed for Cold Terrestrial Exoplanets.” The Astronomical Journal 160 (November): 238. https://doi.org/10.3847/1538-3881/abb8df.
Menou, Kristen. 2012. “Magnetic Scaling Laws for the Atmospheres of Hot Giant Exoplanets.” The Astrophysical Journal 745 (February): 138. https://doi.org/10.1088/0004-637X/745/2/138.
Mikal-Evans, Thomas, Ian J. M. Crossfield, Björn Benneke, et al. 2021. “Transmission Spectroscopy for the Warm Sub-Neptune HD 3167c: Evidence for Molecular Absorption and a Possible High-metallicity Atmosphere.” The Astronomical Journal 161 (January): 18. https://doi.org/10.3847/1538-3881/abc874.
Morley, Caroline V., Heather Knutson, Michael Line, et al. 2017. “Forward and Inverse Modeling of the Emission and Transmission Spectrum of GJ 436b: Investigating Metal Enrichment, Tidal Heating, and Clouds.” The Astronomical Journal 153 (February): 86. https://doi.org/10.3847/1538-3881/153/2/86.
Movshovitz, Naor, Jonathan J. Fortney, Chris Mankovich, Daniel Thorngren, and Ravit Helled. 2020. “Saturn’s Probable Interior: An Exploration of Saturn’s Potential Interior Density Structures.” The Astrophysical Journal 891 (March): 109. https://doi.org/10.3847/1538-4357/ab71ff.
Piaulet, Caroline, Björn Benneke, Jose M. Almenara, et al. 2023. “Evidence for the Volatile-Rich Composition of a 1.5-Earth-radius Planet.” Nature Astronomy 7 (February): 206–22. https://doi.org/10.1038/s41550-022-01835-4.
Sellke, Thomas, M. J Bayarri, and James O Berger. 2001. “Calibration of ρ\rho Values for Testing Precise Null Hypotheses.” The American Statistician 55 (1): 62–71. https://doi.org/10.1198/000313001300339950.
Sing, David K., Zafar Rustamkulov, Daniel P. Thorngren, et al. 2024. “A Warm Neptune’s Methane Reveals Core Mass and Vigorous Atmospheric Mixing.” Nature 630 (8018): 831–35. https://doi.org/10.1038/s41586-024-07395-z.
Teske, Johanna K., Daniel Thorngren, Jonathan J. Fortney, Natalie Hinkel, and John M. Brewer. 2019. “Do Metal-rich Stars Make Metal-rich Planets? New Insights on Giant Planet Formation from Host Star Abundances.” The Astronomical Journal 158 (December): 239. https://doi.org/10.3847/1538-3881/ab4f79.
Thao, Pa Chia, Andrew W. Mann, Adina D. Feinstein, et al. 2024. “The Featherweight Giant: Unraveling the Atmosphere of a 17 Myr Planet with JWST.” The Astronomical Journal 168 (December): 297. https://doi.org/10.3847/1538-3881/ad81d7.
Thorngren, Daniel P. 2024. The Hot Jupiter Radius Anomaly and Stellar Connections. arXiv. https://doi.org/10.48550/ARXIV.2405.05307.
Thorngren, Daniel P., and Jonathan J. Fortney. 2018. “Bayesian Analysis of Hot-Jupiter Radius Anomalies: Evidence for Ohmic Dissipation?” The Astronomical Journal 155 (May): 214. https://doi.org/10.3847/1538-3881/aaba13.
Thorngren, Daniel P., Jonathan J. Fortney, Eric D. Lopez, Travis A. Berger, and Daniel Huber. 2021. “Slow Cooling and Fast Reinflation for Hot Jupiters.” The Astrophysical Journal 909 (March): L16. https://doi.org/10.3847/2041-8213/abe86d.
Thorngren, Daniel P., Jonathan J. Fortney, Ruth A. Murray-Clay, and Eric D. Lopez. 2016. “The Mass-Metallicity Relation for Giant Planets.” The Astrophysical Journal 831 (November): 64. https://doi.org/10.3847/0004-637X/831/1/64.
Thorngren, Daniel P., Eve J. Lee, and Eric D. Lopez. 2023. “Removal of Hot Saturns in Mass-Radius Plane by Runaway Mass Loss.” The Astrophysical Journal 945 (March): L36. https://doi.org/10.3847/2041-8213/acbd35.
Thorngren, Daniel P., David K. Sing, and Sagnick Mukherjee. 2025. Bayesian Model Comparison and Significance: Widespread Errors and How to Correct Them. arXiv. https://doi.org/10.48550/arXiv.2510.00169.
Thorngren, Daniel, and Jonathan J. Fortney. 2019. “Connecting Giant Planet Atmosphere and Interior Modeling: Constraints on Atmospheric Metal Enrichment.” The Astrophysical Journal 874 (2): L31. https://doi.org/10.3847/2041-8213/ab1137.
Thorngren, Daniel, Peter Gao, and Jonathan J. Fortney. 2019. “The Intrinsic Temperature and Radiative-Convective Boundary Depth in the Atmospheres of Hot Jupiters.” The Astrophysical Journal 884 (October): L6. https://doi.org/10.3847/2041-8213/ab43d0.
Tremblin, P., G. Chabrier, N. J. Mayne, et al. 2017. “Advection of Potential Temperature in the Atmosphere of Irradiated Exoplanets: A Robust Mechanism to Explain Radius Inflation.” The Astrophysical Journal 841 (May): 30. https://doi.org/10.3847/1538-4357/aa6e57.
Valsecchi, Francesca, Frederic A. Rasio, and Jason H. Steffen. 2014. “From Hot Jupiters to Super-Earths via Roche Lobe Overflow.” The Astrophysical Journal 793 (1): L3. https://doi.org/10.1088/2041-8205/793/1/L3.
Vissapragada, Shreyas, Michael Greklek-McKeon, Dion Linssen, et al. 2024. “Helium in the Extended Atmosphere of the Warm Superpuff TOI-1420b.” The Astronomical Journal 167 (May): 199. https://doi.org/10.3847/1538-3881/ad3241.
Wahl, Sean M., Daniel Thorngren, Tiger Lu, and Burkhard Militzer. 2021. “Tidal Response and Shape of Hot Jupiters.” The Astrophysical Journal 921 (November): 105. https://doi.org/10.3847/1538-4357/ac1a72.
Yadav, Rakesh K., and Daniel P. Thorngren. 2017. “Estimating the Magnetic Field Strength in Hot Jupiters.” The Astrophysical Journal Letters 849 (November): L12. https://doi.org/10.3847/2041-8213/aa93fd.
Yee, Samuel W., Gudmundur Stefánsson, Daniel Thorngren, et al. 2025. “The Super-puff WASP-193 b Is on a Well-aligned Orbit.” The Astronomical Journal 169 (April): 225. https://doi.org/10.3847/1538-3881/adba5f.
Yoshida, Stephanie, Shreyas Vissapragada, David W. Latham, et al. 2023. TESS Spots a Super-puff: The Remarkably Low Density of TOI-1420b.” The Astronomical Journal 166 (November): 181. https://doi.org/10.3847/1538-3881/acf858.